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Townsend's Vole
If you live west of the Cascades in Washington and something is eating your lawn, your bulbs, or the roots of your young fruit trees, this is the animal Washington State University Extension names first. Its vole publication says it in one line: the two species most often responsible for vole-related backyard and garden damage in Washington are Townsend's vole west of the Cascade Mountains and the montane vole east of them. That one sentence is the whole reason it matters to keep this animal separate from the creeping vole. The creeping vole is a forest and clearcut animal whose documented pest role is girdling conifer seedlings on reforestation ground. Townsend's vole is the yard animal. It works lawns, gardens, orchards, cropped ground, pasture, wet meadow and marsh edge, and Oregon State University Extension gives it a second name that tells you exactly where to look for it: bog rat. Three things set it apart. It is the largest of the west-side lowland garden voles, up to about 9.5 inches. It attains the highest average density of any Microtus in North America, with documented cyclic peaks near 700 animals per hectare. And it nests above the waterline, on hummocks, which is what lets it hold soggy winter ground that would drown out its competitors. One more thing before you read further, because it catches researchers as well as homeowners: this is not Townsend's chipmunk. Different family entirely. They share a name because they share a namesake.

Seasonal Timeline

Townsend's voles do not hibernate and are active in every month of the year, day and night. What changes with the season is where the damage lands. Green grass and forbs carry them through the growing season, and in winter, when other food is scarce, they turn to bulbs, roots and the bark of woody plants. That is when bulb beds get emptied and young trees get girdled. The runway network in a lawn is usually most visible in late winter and early spring, when the grass is flat and the worn trails are exposed.

Breeding seasonMar–Sep
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Townsend's voles do not hibernate and are active in every month of the year, day and night. What changes with the season is where the…

Nothing about this animal shuts down for winter, and the winter is when it costs you money. Washington State University Extension states that these voles prefer succulent grasses, forbs, roots and bulbs but will readily feed on the bark and roots of woody plants during winter when other food sources are scarce. Its Hortsense fact sheet is blunter: bulbs and roots are often completely eaten over winter, and trees can be completely girdled. California's own wildlife habitat account records the species as active year-round, mostly nocturnal but sometimes active throughout the day, and the species monograph agrees that it is active both day and night.

Breeding has two versions in the record, and neither should be blended into the other. California's account gives breeding primarily from March to September, with year-round breeding possible in lowland areas and at least two litters a year, noting that large voles are able to breed in winter. The species monograph makes a more interesting point: the length of the breeding season in this species is tied to where the population sits in its multiannual abundance cycle. On one British Columbia island, breeding ran from May to October in one year, from early February to mid-August the next, and from mid-February to May the year after that. There is no fixed calendar to give, and the monograph's position is that there should not be one.

The 3 to 5 year cycle is the number a homeowner should actually plan around. Washington State University records periodic vole population explosions in the open grassland, field and forest-edge ground that borders yards, commonly occurring every 3 to 5 years, with populations that may number more than 1,000 voles per acre before crashing. The research literature backs the scale: a review of long-running British Columbia populations identified four probable cycles with peaks between 525 and 800 voles per hectare, averaging 697. A quiet year is not evidence the problem has gone away.

Where They're Found

This is a west-side, low-elevation, wet-ground animal, and its eastern boundary is the point. The documented range runs from Triangle Island, British Columbia south to Humboldt Bay, California, reaching east in British Columbia only to Chilliwack, in Washington to Sauk, Nisqually Flats and Clark County, and in Oregon to Salem, Eugene and Prospect. That is the Willamette Valley floor and the coastal lowlands, not the Cascade crest. Altitudinally the species runs from sea level to about 1,830 meters, roughly 6,000 feet, in the Olympic Mountains, and to about 914 meters, roughly 3,000 feet, in the Oregon Cascades.

The Pacific Northwest is the core of its range. Washington State University Extension names Townsend's vole as the west-of-the-Cascades half of the pair of species most often responsible for backyard and garden damage in Washington, and lists its habitat as lawns, gardens, orchards, cropped areas, open grasslands, farms, meadows and young forested plantations. Oregon State University Extension places it primarily on the coast and in the Coast Range, in meadows, marshy areas, pastures and riparian areas with dense ground cover. Older survey work fills in the picture: the nominate subspecies lives in marshes or damp meadows under deep rank vegetation and avoids forested areas, while along the Oregon coast it occupies deflation plains, willow and sedge marshes, wet pasturelands and tidal river habitats. In the Willamette Valley it has been called chiefly riparian, but it also holds moist meadows and irrigated fields with dense sedge, bracken fern and wild rye.

The California range rests on a state agency source, and it is a narrow one. The California Department of Fish and Wildlife's own life history account states that in California this species is found in coastal Humboldt and Del Norte counties, and that it is abundant in valley foothill riparian, wet meadow and pasture habitats and common in fresh and saline emergent wetlands, croplands, grasslands and early successional forest stages. The species monograph independently puts the southern range limit at Humboldt Bay and Eureka. That is a two-county coastal foothold, not a statewide presence, and it should not be read as putting this animal anywhere near the Central Valley or southern California.

The San Juan Islands hold a named subspecies with its own conservation status. The Shaw Island Townsend's vole is the subspecies Microtus townsendii pugeti, found only in the San Juan Archipelago. Washington's Department of Fish and Wildlife reports confirmed populations on Lopez, Orcas, San Juan and North Finger islands and describes them as robust where they occur, and lists the subspecies as a Species of Greatest Conservation Need under the state's wildlife action plan. That is a planning designation rather than a threatened or endangered listing, and no take prohibition attaches to it, but it is worth knowing before doing anything in the islands. On the ground, that subspecies occupies meadows, salt marshes, driftwood on sea beaches, rock piles and sparse dry grass.

Two states are outside the range. No source places this species in Alaska. Its northern limit is Triangle Island, off the north end of Vancouver Island, with the entire British Columbia coast between there and the Alaska border. Hawaii has no vole of any kind. There is also no fossil record for this species at all, which is unusual enough to say out loud.

On a property, look at the wet, grassy, low ground. Dense herbaceous growth over soft soil is what this animal needs, and it takes the lawn edge, the unmowed margin, the ditch bank, the pasture and the bed with thick mulch over the bare, dry, or forested parts of a yard every time.

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